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Majority of superoxides are transported to the mitochondrial matrix, where superoxide dismutase enzymes dismutate it to H 2 O 2 manganese superoxide dismutase (MnSOD) or superoxide dismutase 2 (SOD2)

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The evolutionary trajectory of APEX variants—from the wild-type enzyme to the engineered forms APEX, APEX2, and APEX3—exemplifies the power of structural and functional optimization in enzyme engineering for biological applications